| 1978 | B.A., Biology/Psychology | Reed College |
| 1985 | Ph.D., Neurobiology | UCSF School of Medicine |
| 1985-87 | Postdoctoral Associate | Columbia University |
| 1987 | Assistant Instructor, Molecular Embryology of the Mouse | Cold Springs Harbor Laboratory |
| 1987-1990 | Postdoctoral Research Associate/Instructor | Tufts University School of Medicine |
| 1991-1995 | Assistant Professor, Department of Biological Sciences | University of Iowa |
| 1995-1996 | Assistant Professor, Anatomy & Neurobiology, Development & Cell Biology | University of California, Irvine |
| 1996-2005 | Associate Professor, Anatomy & Neurobiology, Developmental & Cell Biology | University of California, Irvine |
| 2000 |
Track Director: Molecular Neurobiology Track Graduate Program in Molecular Biology, Genetics and Biochemistry |
University of California, Irvine |
| 2005-Present | Professor, Anatomy & Neurobiology, Developmental & Cell Biology | University of California, Irvine |