| 1983 | M.D. | Stanford University |
| 1987 | Neurology Residency | Stanford University |
| 1991 | Ph.D., Neuroscience | Stanford University |
| 1991-996 | Assistant Professor, Neurology, Anatomy & Neurobiology, Neurobiology & Behavior | University of California, Irvine |
| 1996-2002 | Associate Professor, Neurology, Anatomy & Neurobiology Neurobiology & Behavior |
University of California, Irvine |
| 2002-Present | Professor, Neurology, Anatomy & Neurobiology | University of California, Irvine |